Bioquatic Photo contains a large selection of biological and nature related photos.
Many have been used in various projects. Below are some examples.


     TROPICAL FISH Coloring Book  (2010)

 Can you color the Regal Angelfish?


Sole photo contributor in Sierra Club's TROPICAL Fish Coloring Book, published by Pomegranate Publications Inc., CA, USA (2010).

A total of 22 of my fish photos were used.  Click the name of the fish below to view them online on Bioquatic Photo's Web Site:

Regal Angelfish   Picasso Fish   Copperband Butterflyfish   Mandarin Dragonet   Harlequin Filefish   Red Lionfish   Yellowtail Coris   Emperor Angelfish   Clown Triggerfish   Ornate Cowfish   Spotted Sea Horse   Flame Angelfish   Sunset Wrasse   Seven-Stripe Frontosa   Klunzinger's Wrasse   Blue-Girdled Angelfish   Zebra Turkeyfish      Common Discus   Queen Triggerfish   Threadfin Butterflyfish   Harlequin Tuskfish



 WHAT FISH?  (2009)  

Major photo contributor in Phil Hunt's book WHAT FISH? A Buyer's Guide to Reef Fish published by INTERPET Publishing, Dorking, Surrey, UK (2009).

ISBN: 978 1 84286 222 3

A total of 58 of my photos were used, including the ones shown here (click the image numbers to learn more about these images):

2637-D: Yellow Watchman gobies (Cryptocentrus cinctus) with symbiotic shrimp (Alpheus sp.)
2208: Bicolor blenny (Ecsenius bicolor)
2210: Red Sea Mimic Blenny (Ecsenius gravieri)
2629-D: Filamented-finned Prawn Goby (Stonogobiops nematoda)
35058-D and 35086-D: Yasha Hase Shrimp Goby (Stonogobiops yasha)




Q&A The Marine Aquarium  (2008)

Major photo contributor in Dr Ashley Ward's book Q&A Questions and Answers The Marine Aquarium published by INTERPET Publishing, Dorking, Surrey, UK (2008).

ISBN: 978-1-84286-168-4

Photos of invertebrates are numerous in Bioquatic's collection. A total of 52 of my photos were used, including the ones shown here (click the image numbers to learn more about these images):

2211: Social feeding group of the Poweder Blue Tang (Acanthurus leucosternon) from a shallow reef in the Maldives (A rare image and one of my most used images shot in 1980 when the Maldives were totally unspoiled)
35545: Bigeye Trevally (Caranx sexfasciatus) from the Coral Sea






Major photo contributor in Tristan Lougher's book WHAT INVERTEBRATE? A Buyer's Guide for Marine Aquariums published by INTERPET Publishing, Dorking, Surrey, UK (2007).

ISBN: 978 1 84286 179 0

Photos of invertebrates are numerous in Bioquatic's collection. A total of 111 of my photos were used, including the ones shown here (click the image numbers to learn more about these images):

5598: Staghorn Corals (Acropora formosa and Acropora nobilis) from the Great Barrier Reef
8385-D: Blue knuckle Hermit Crab (Calcinus elegans)
8317-D: Cave Cleaner Shrimp (Urocardiella antonbruunii)





Major photo contributor in Matthew L. Wittenricht's book The Complete Illustrated Breeder's Guide to Marine Aquarium Fishes published by T.F.H. Publications (NJ, USA) and Microcosm Ltd., (VT, USA), (2007).

ISBN: 1-890087-71-9

A total of 48 of my photos were used, including the ones shown here (click the image numbers to learn more about these images):

35137 and 35134 showing male and female of the rare Resplendent Pygmy Angelfish (Centropyge resplendens) endemic to Ascension Island in the central Atlantic.




WHAT FISH?  (2006)


Major photo contributor in Tristan Lougher's book WHAT FISH? A Buyer's Guide to Marine Fish published by INTERPET Publishing, Dorking, Surrey, UK (2006).

ISBN: 978 1 84286 118 9

A total of 21 of my photos were used, including the ones shown here (click the image numbers to learn more about these images):

2659-D: Multibanned Goby (Elacatinus multifasciatus)
2523: Peppermint Basslet or Swissguard Basslet (Liopropoma rubre)



 500 Ways to be a better Marine Fishkeeper  (2005)

Major photo contributor in Dave Garratt, Tim Hayes, Tristan Lougher and Dick Mills' book 500 Ways to be a better MARINE FISHKEEPER published by INTERPET Publishing, Dorking, Surrey, UK (2005).

ISBN: 1-84286-108-5

A total of 66 of my photos were used, including the ones shown here (click the image numbers to learn more about these images):

8080-D: Mithrax crab (Mithrax sculptus)
8310: White-patched anemone shrimp (Periclimenes brevicarpalis)
2245: Citron Goby (Gobiodon citrinus)




Kirken ved Havet  (2004)


Major photo contributor in Inger Vågen's (editor) Kirken ved Havet (freely translated to "The Church by the Sea"), published by Hidra Menighet (Hidra, Norway), (2004).

ISBN: 82-303-0174-3

A total of 32 of my photos were used, including the one shown bewlow (click the image number to learn more about this image):

71019 showing Hidra Kirke by night.

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By Alf Jacob Nilsen and Svein A. Fosså


Languages:     Trilingual English / French / Dutch
               90-76886-09-1 E / F / D
           240 x 265 mm
176 pages
               Alf Jacob Nilsen and Svein A. Fosså
Various photographers
         About 240 images full colour
Semi-mat art paper 150 gr
Linen hard bound
     Glossy finish (full colour)

Publisher:       Tectum Publishers and
                       Loft Publications, Antwerp, Belgium.

"an amazon dreamscape" Photo:Mike Duhon
Tank built and maintained by Aquarium Design Group, Texas, USA
Recent aquarium books are more beautiful and better equipped than ever. The factual contents have a quality unforeseen. Fishes, invertebrates and plants are identified with scientific accuracy. Well-educated and serious professionals describe the technical equipments, foods and maintenance routines with dedication and seriousness. The faith of the aquarium animals has never been in better hands. The likeliness to achieve success, as an aquarium keeper, is greater today than ever before.  
"builders of coral reefs" Photo: Werner Huthmacher
Tank constructed and owned by Pieter Van Suijlekom, the Nederland
Still, to a large extent, modern aquarium books have lost track of one of the major considerations people have when contemplating setting up an aquarium in the first place. Surely, quite a few new aquarists have chosen their hobby foremost in order to study the aquatic world really up-close, but in a modern home, more and more people are concerned about what the aquarium can do to the interior of their own home.  
"in the deep of the blue" Photo: Werner Huthmacher

In Tectum's newest book project "The Most Beautiful Aquariums of the World" the sole focus is, for the first time in any modern aquarium book, on what an aquarium can do to your home. How a well-designed aquarium can become a focal point in even the most fashionable of living rooms. The Tectum team selected and photographed the top-of-the-top aquariums from amongst several of the most discriminating aquarium owners in the world.

“The Most Beautiful Aquariums of the World” is not your average aquarium book. It is a book about the use of aquariums as decorative elements and focal points in room furnishing. This book focuses at architecture, decoration, furniture, as well as aquariums and living aquatic organisms. Through sequences of beautiful photography and short pieces of texts, it tells the story of how a small segment of nature, in the form of life in a glass encased box, can be a part of a beautiful room interior.

"the amazon"
Photo and aquarium construction by: Reef Aquaria Design Inc., Director Jeff Turner, Coconut Creek, Florida, USA, a major contributor to the content of the book..



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Korallenriff-Aquarium - Band 1: Grundlagen für den erfolgreichen Betrieb

ISBN: 978-3-86659-072-4




S.A. Fosså und A. J. Nilsen

  • 480 Seiten
  • 587 Farbfotos, Zeichnungen, Tabellen, Grafiken
  • Format: 17,5 x 23,2 cm, Hardcover


Das Korallenriff-Aquarium – Band 1

Die sechsbändige Reihe „Das Korallenriff-Aquarium“ hat im Hobby Maßstäbe gesetzt. Im hier vorliegenden, völlig neu bearbeiteten, erweiterten und aktualisierten Band 1 erläutern die Autoren die Grundlagen, die zur erfolgreichen Riffaquaristik nötig sind. Sie schildern die Entstehung sowie die verschiedenen Typen von Riffen und zeigen, wie ein herrliches, artenreiches Korallenriff  im Aquarium geschaffen und gepflegt werden kann.

Fundierte, leicht verständliche Information und brillante Fotos: Das unentbehrliche Standardwerk für jeden Meerwasser-aquarianer!


Die verschiedenen Rifftypen und ihre Ökologie
Das Korallenriff-Aquarium:

  • Beleuchtungssysteme in der Praxis
  • Das optimale Meerwasser: Worauf ist zu achten?
  • Stoffkreisläufe: Stickstoff und Phosphor messen und kontrollieren
  • Konstante Temperaturen: Heiz- und Kühlsysteme
  • Filterung und Strömung: Welche Geräte wirklich nötig sind
  • Ausgewählte Porträts sehenswerter Aquarien



 1) Korallenriffe
Aspekte zur geologischen Geschichte der Korallenriffe

Typen von Korallenriffen
  • Kaltwasser-Tiefseeriffe
  • Tropische Korallenriffe

Geografische Verbreitung tropischer Korallenriffe
Bedrohungen für die Korallenriffe

  • Die Korallenbleiche (Coral Bleaching)
  • Verminderte Kalzifizierungsrate
  • Korallenkrankheiten
  • Dornenkronenseestern (Crown of Thorns Starfish, COTS)
  • Sedimenteinspülungen und erhöhte Nährstoffkonzentration

Einführung in die Ökologie der Korallenriffe

  • Energie
  • Primärproduzenten, Konsumenten und abbauende Organismen
  • Diversität

Allgemeine Korallenriffökologie

  • Fotosynthese
  • Respiration (Zellatmung)
  • Produzenten, Konsumenten und abbauende Organismen
  • Anorganische Nährstoffe im Korallenriff
  • Organische Substanzen in Korallenriffen

Primärproduktion im Korallenriff
Primärproduzenten im Korallenriff
Konsumenten im Korallenriff
Mikroorganismen des Korallenriffs
Die Zonen des Korallenriffs und der Besuch einiger spezifischer Riffe

  • Ein Saumriff im Roten Meer
  • Das Große Barriereriff
  • Ein Saumriff bei Nananu-I-Ra, Fidschi
  • Ein „Algenriff“ von South Beach, Rarotonga, Cook Island
2) Ein Korallenriff im Aquarium
Wie alles begann

Die Aquaristik erregt öffentliches Interesse
  • Die Ankunft exotischer Arten

Moderne Versorgung mit Riffbewohnern

  • Die Vertriebskette
  • Wildfänge
  • Vermehrung in Menschenhand
  • Das Exportgeschäft

Pflichten des Hobbyaquarianers

  • Fressgewohnheiten
  • Aggressives Verteidigungs- und Fressverhalten
  • Gifte
  • Empfindlichkeit
  • Stabilität – eine Frage der Geduld
3) Licht
UV-Strahlung und ihre Wirkung auf Korallen

Korallenfarben und Fluoreszenzpigmente
  • Anregung und Emission
  • Pigmentverbindungen
  • Biologische Funktionen von Fluoreszenzproteinen

 Licht und Fotosynthese
 Beleuchtungsstärke und die Lichtverhältnisse im Meer
 Bedeutung des Lichts für das Korallenriff-Aquarium
 Lichtquellen für das Korallenriff-Aquarium

  • Leuchtstofflampen
  • T5 und T8
  • Aktinisch blaue Leuchtstoffröhren
  • Kombination von Leuchtstofflampen
  • Halogen-Metalldampflampen (HQI)
  • LED-Beleuchtung
 Einige praktische Tipps und Hinweise

4) Wasser
Salzgehalt des Meerwassers

            - Das Messen der Salinität
Osmolarität und osmotische Regulation
Künstliches oder natürliches Meerwasser?
Chemische Reaktionen und Parameter des Meerwassers
  1. Der pH-Wert
  2. Pufferkapazität und Puffervermögen
  3. Alkalinität und Karbonathärte
  4. Kalzium
  5. Kohlendioxid
  6. Magnesium
  7. Strontium
  8. Andere Haupt-Ionen des Meerwassers
  9. Sauerstoff
  10. Abbauprodukte
  11. Silikate
  12. Spurenelemente

5) Temperatur

  • Was ist Korallenbleichen?
  • Vorkommen von Korallenbleichen
  • Warum taucht das Phänomen „Korallenbleichen“ zurzeit auf und was müssen wir in Zukunft erwarten?

 Beheizung des Korallenriff-Aquariums
 Kühlung des Korallenriff-Aquariums

  • Notfallmaßnahmen bei Überhitzung
6) Biochemische Prozesse im Korallenriff-Aquarium
Stickstoff und Stickstoffzyklus

Bakterien und ihr Milieu
Der Stickstoffzyklus – detailliert betrachtet
  1. Mineralisation oder Ammonifikation
  2. Nitrifikation
  3. Assimilation oder Nitratreduktion
  4. Denitrifikation

Phosphat und der Phosphorzyklus


7) Filterung, Wasserpflege und Strömungstechnik
Biologische Filterung

  1. Biofilter mit Lebenden Steinen
  2. Biofilter mit anderen Filtermaterialien
  3. Rieselfilter
  4. Denitrifikationsfilter
  5. Monokulturfilter
  6. Sandbett und Sandbettfilter
  7. Mechanische Filter

Kohlefilter und andere chemische Adsorptionsfilter

  • Phosphatfilter

Wasserbewegung und Strömungstechnik
Kombination von Filtern


8) Ausgewählte Aquarienporträts


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A fresh, easy-to-follow approach to setting up an strcking a reef aquarium,
filled with simple but authoritative advises


Reef Secrets includes:
  • A profusely illustrated guide to selecting fishes, corals, and other invertebrates.
  • Lessons from the wild to create simpler, more stable reef aquariums.
  • Secrets of good water quality, water motion, lighting, aquascaping, live rock, and substrates.
  • Understanding algae, feeding, reef ecology. 
  • Suggestions for unique biotope reef systems.

Authors: Alf Jacob Nilsen & Svein A. Fosså
Publisher: Microcosm / TFH Publications
ISBN: 189008767X
Product Dimensions: 11.2 x 8.6 x 0.8 inches
Hardcover: 240 pages

  1. Light, Darkness and Shadows 
  2. Water Quelities & MOvements
  3. Aquascaping
  4. Biotopes
  5. Aquarium Stocking
  6. Stocking Guide
  7. Reef Algae
  8. Foods & Feeding
  9. Friends of the Reef
Stocking Guide

"A Consice Encyclopedia of Fishes and Invertebrates Commonly Available to the Marine Aquarist"

The stocking guide covers all major phyla and 440 of the most common organisms kept in captivity. 

The guide gives short, but precise information on: Systematic, Scientific- and popular names, Distribution, Lighting, Circulation, Water Quality, Aquascaping, Feeding, Compatibility, Hazards and Conservation.


Best marine aquarium book ever, February 2, 2004

"This book is great, it is not only a book packed with usful infomation for anyone beginer or advanced, it has a lot of info on what equiptement you need and tells you why you need it. It also is just a great book on hand. I would reccomend it to everyone".   

UK Only' Aquarist Classifieds Readers Review: 

An ideal introduction to reefkeeping. This book is an ideal introduction to reefkeeping, covering the basic techniques and accepted best practice with a chapter each on lighting requirements, maintaining water quality, aquascaping, algae, feeding, and conservation. There is a stocking guide covering around 400 of the more commonly available species of fish, corals and invertebrates. Each brief species description is accompanied by an excellent photo and clearly set out advice on their care requirements and the degree of challenge. For me though, what made this book stand out from many others covering similar territory were the two chapters on modelling and stocking aquariums to try to emulate real biotopes and ecological niches. They provide inspirational, yet practical, examples of the different sorts of themes you could consider - be it a Red Sea cave or a reef gorge - and how to implement them. This has been done before, most notably in John Tullock's 'Natural Reef Aquariums', but that just did not affect me the way this book has. Nor is Tullock's book as current - a real issue given the speed with which this hobby is evolving.

Nilsen and Fossa know what they are talking about. They are the authors of the four volume Modern Coral Reef Aquarium series of books - probably the most comprehensive and respected European reference books on reefkeeping and the identification and care of the species inhabiting them. So in summary: a well written, beautifully illustrated, comprehensive and inspirational guide to reefkeeping by two authoritative and internationally respected authors.

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The Modern Coral Reef Aquarium Vol. 4

Aquarium keeping of snails, mussels, sea stars, brittle stars, sea urchins, sea cucumbers, 
tunicates, and more!

The Modern Coral Reef Aquarium Vol. 4 covers several important invertebrate groups of great interest and importance to the modern reef aquarium keeper: Molluscs, Echinoderms, Tunicates and more!

Hardcover, 480 pages, 1030 color photos, 60+ line drawings and B/W photos, several tables. 

ISBN 3-928819-22-4.
Birgit Schmettkamp Verlag, Bornheim, Germany.


  1. The Future of Reef Aquariums
  2. Mollusca
  3. Echinodermata
  4. Tunicates and Lancelates

Molluscs, such as snails, slugs and bivalves, commonly found in the trade are discussed in detail. Like in previous volumes of this book series, also more obscure and rare members of the phylum are covered to the extent, which can be of value to the marine aquarist.

Echinoderms - or spiny skinned animals - are shown in spectacular photos, and the biology and aquarium care of the many interesting brittle stars, starfishes, sea cucumbers, and sea urchins are described in detail.


e book also discusses the interesting group of sea squirts, or tunicates. These organisms, which are very insufficiently covered in most aquarium books, are dealt with in detailed accuracy and illustrated with magnificent colour photos.  

The world number one expert, the late Dr. Patricia Kott of Queensland Museum, Brisbane, Australia has contributed greately to make this chapter one of the most comprehensive overviews of tropical tunicates!

As an introduction to the book, Dr. Bruce Carlson (former Director of the Waikiki Aquarium, Hawaii) has written a fascinating chapter on "The Future of Reef Aquariums".

Together with the other three MCRA volumes, this book completes your knowledge on the invertebrates of the "Modern Coral Reef Aquarium"!

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